HE400 feels generally dynamically looser and more V-shaped. HE4 has a very grainless timbre save for a little distortion spike at 5k giving a perception of 10k presence. HE4 tone is more like the 5LE. Better resolve, transients, and staging on the HE4. HE400 I could never get quite perfect with mods, but close. My modded HE4 is smooth as butter and slightly dark, but great extension on either end.
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